Time-lapse of a wild-type embryo, visualized with an actin marker (LIFEACT::GFP, left) expressed in the epidermis and a nuclear marker (HIS-24::mCHERRY, right) expressed in muscles, showing displacement of actin filaments and muscle nuclei due to muscle contractions (Zhang et al.,2011), and rolling movements. The embryo is 10 min past the 2-fold stage (80 min after ventral enclosure). The movie on the right was started around 20 s after the movie on the left.