To view this video please enable JavaScript, and consider upgrading to a web browser that supports HTML5 video Movie 2 Direction of rotation opposite to the first harmonic ellipse. EFA mode 3 generating a shape with 4 features, its rotation direction opposite to the rotation direction of the first harmonic. Morphometrics of complex cell shapes: lobe contribution elliptic Fourier analysis (LOCO-EFA) Yara E. Sánchez-Corrales, Matthew Hartley, Jop van Rooij, Athanasius F.M. Marée, and Verônica A. Grieneisen Development 2018. 145:None-None; doi: 10.1242/dev.156778 Download Original Video