To view this video please enable JavaScript, and consider upgrading to a web browser that supports HTML5 video Movie 3 Opening reaction of the inner valve lids of the mesothoracic spiracle on a tactile stimulus (filter trichomes are partly removed). The opening is quicker than the closing. Opening starts from ventral and closing starts from dorsal. Structure of the thoracic spiracular valves and their contribution to unidirectional gas exchange in flying blowflies Calliphora vicina Lutz T. Wasserthal, and Anja S. Fröhlich Journal of Experimental Biology 2017. 220:208-219; doi: 10.1242/jeb.149013 Download Original Video