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Nocturnal bigeye (Pempherus adspersa) in their daytime, reef-crevice refuge provide a strong theoretical case for vestibular cues in rheotaxis.

Rheotaxis revisited: a multi-behavioral and multisensory perspective on how fish orient to flow

Sheryl Coombs, Joe Bak-Coleman, and John Montgomery

Journal of Experimental Biology 2020. 223:None-None; doi: 10.1242/jeb.223008