S. polymorpha and S. sesxpinosus locomoting on flat, less rugose, and more rugose terrain. S. polymorpha locomoting on (A) flat (Rg = 0), (B) less rugose (Rg = 0:17), and (C) more rugose (Rg = 0:44) terrain. S. sexspinosus locomoting on (A) flat (Rg = 0), (B-D) less rugose (Rg = 0:17), and (E) more rugose (Rg = 0:44) terrain. All videos playback at real-time and 0.1x speed.